He starred as cop killer Barry Weiss in the British crime-thriller Blitz and in the British horror film Wake Wood. He appeared in seven seasons, until his character's death in the season 7 finale " The Dragon and the Wolf".
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In 2011, Gillen began playing Petyr "Littlefinger" Baelish on the HBO series Game of Thrones, for which he received his second Irish Film & Television Award nomination. He co-starred as Phil Hendrick in the British drama Thorne. He appeared in the 2009 film 12 Rounds, and in July of that year, he appeared in the one-off BBC2 drama Freefall. In 2008, he was named an "Irish cult hero" by the Sunday Tribune. In 2004, having been spotted by producers in The Caretaker, Gillen was cast as Tommy Carcetti in the HBO series The Wire, for which he received an Irish Film & Television Award for Best Actor in a Lead Role in Television. He was nominated for a Tony Award for his Broadway role in Harold Pinter's play The Caretaker and has also been nominated for an Irish Times Theatre Award for his portrayal of Teach in the Dublin Gate Theatre's 2007 production of David Mamet's American Buffalo. Gillen played Stuart Alan Jones in the Channel 4 television series Queer as Folk and its sequel, for which he received a British Academy Television Award nomination for Best Actor. He moved to London in 1987 when he was nineteen. The name Aidan Murphy was already registered so he began using his mother's maiden name as a stage name. Gillen began his acting career as a teenager, joining the National Youth Theatre at the age of fourteen and playing Nick Bottom in a production of A Midsummer Night's Dream at the Project Arts Centre when he was sixteen. Gillen was born Aidan Murphy in the Drumcondra area of Dublin on 24 April 1968, the youngest of six children born to Patricia (née Gillen) and Denis Murphy. Gillen has won three Irish Film & Television Awards and has been nominated for a British Academy Television Award, a British Independent Film Award, and a Tony Award.
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In 2021, he had a role in the dramatic crime series Mayor of Kingstown. He also provided the voice and motion capture for Paul Serene in the 2016 video game Quantum Break. Allen Hynek in The History Channel's Project Blue Book (2019–2020). He is known for his film and television roles including Stuart Alan Jones in the Channel 4 series Queer as Folk (1999–2000), Tommy Carcetti in the HBO series The Wire (2004–2008), John Boy in the RTÉ series Love/Hate (2010–2011), Petyr "Littlefinger" Baelish in the HBO series Game of Thrones (2011–2017), CIA operative Bill Wilson in The Dark Knight Rises (2012), Janson in Maze Runner: The Scorch Trials (2015) and Maze Runner: The Death Cure (2018), Aberama Gold in the BBC One series Peaky Blinders (2017–2019), Queen's manager John Reid in Bohemian Rhapsody (2018), and Dr. Aidan Murphy (born 24 April 1968), better known as Aidan Gillen ( / ˈ ɡ ɪ l ə n/, Irish: Aodhán Ó Giolláin), is an Irish actor.